Trachyplax arctica, a new multiplated problematic fossil from the Lower Cambrian of North Greenland

A new scleritome−bearing organism with eight sclerite types, Trachyplax arctica gen. et sp. nov., is described from the lower Cambrian Paralledal Formation of North Greenland. The originally calcareous sclerites are now silicified; no microstructures are preserved. The dominant sclerite type (A; maximum dimension 19.3 mm) is bilaterally symmetrical, strongly arched, with an oval shield showing co−marginal growth lines and a projecting rostrum with prominent radial ornamentation. A similar sclerite morphology can be identified in Silurian–Carboniferous multiplacophoran molluscs but the remaining sclerite types, which also display a combination of concentric and radial ornamentation, find no clear equivalents. Two models for scleritome reconstruction are presented, based on the relative abundance of the sclerites, but neither promotes a satisfactory assignment to a higher taxon. Despite the morphological dissimilarities, possibly reflecting the age discrepancy, reference to the Multiplacophora is most attractive and entails a substantial extension of the known geological range of that group.