A comparative study of male genitalia in Opatrinae sensu Medvedev [1968] [Coleoptera: Tenebrionidae], with notes on the tribal classification. Part I

A total of 180 species (including 62 type-species of genera) of 111 genera in all the tribes and subtribes included in the Opatrinae sensu Medvedev (1968) [=Opatrini sensu Doyen and Tschinkel (1982)] have been examined. The terminology of the male genitalia within the taxon has been standardized. The use of the aedeagus structure in the classification of Tenebrionidae is discussed. Four main types of aedeagus structure in the Opatrinae have been distinguished: opatrinoid, dendaroid, platynoid and oncotinoid, as well as three additional groups of genera (blapstinoid, melanimoid and dissonomoid), whose appurtenance to the Opatrinae are debatable and requires verification.